Grizzly bears, grey wolves, bison, pronghorn, elk, moose, and river otters were extirpated from the park area in the 19th and early 20thcenturies. In 1913 and 1914, elk from Yellowstone National Park were transported and released into the area that would become Rocky Mountain National Park by the Estes Park Protective and Improvement Association and the USFS. Without large predators, elks thrived but quickly overgrazed vegetation in the park so they are now maintained at more sustainable numbers. In 1978, the Colorado Division of Wildlife reintroduced moose in North Park and river otters in Rocky Mountain National Park. Both species are now frequently seen in Rocky Mountain National Park, particularly on the west side.